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Pair-wise interactions have been observed to promote social hierarchies within groups of animals where individuals with successful agonistic behaviors often achieve dominance. Citation needed Engaging in agonistic behavior can be very costly and thus there are many examples in nature of animals who achieve dominance in more passive ways. Page needed a b c d e f g h Huntingford, Felicity, and Angela. Citation needed In eusocial species, decrease of fertility is among the main reasons for ranking displacement. Laboratory experiments have shown that when foundresses are injected with juvenile hormone (JH a hormone responsible for regulating growth and development in many insect species including wasps, the foundresses exhibit an increase in dominant behavior compared to those who had not been injected. Since nuptial flights are seasonal and workers are wingless, workers are almost always virgin, and (as gamergate ants or laying worker bees ) can only lay unfertilised eggs. This is most likely a function of two factors.

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